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NCOA4, as a key receptor for ferritinophagy, can specifically bind to the conserved surface arginine residues of the ferritin heavy chain at its C-terminus, promoting iron release during autophagosome formation and autolysosome maturation (Liu et al., 2025)

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If the ligand is dimeric, the dimeric fusion may bind to the dimeric ligand bivalently, conferring binding avidity: namely, binding strength is enhanced